在1999年,一艘外星宇宙战舰坠落到了地球,人们用了了十年的时间修复并取名为"MACROSS"。2009年,MACROSS要塞正在举办一场偶像歌星林明美(饭岛真理 配音)的演唱会,突然空袭警报响起,要塞开始了变形。战斗中,一条辉(长谷有洋 配音)驾驶着变形战斗救下一名少女,战机却由于速度过快撞进外层舱室。安定后,辉发现救得是林明美。俩人相识后,为使明美高兴,辉偷开教练机带着明美去兜风。敌人趁机出现,好在朋友这时挺身相救,经过一场混战,辉和早濑未沙(土井美加 配音)醒过来,发现在一个荒芜的星球上。他们了解到,五十万年前,一个叫"普罗多"的古代文明发展到顶盛时,已经可以借遗传密码无性繁殖后代,于是国家分裂为由男性组成的"杰特拉帝"和由女性组成的"赫而特兰帝"开始了长达数万年的战争。在战争中文明急剧衰退了,曾经伟大的文化被遗忘殆尽,退化成了只会战斗的种族,为了维护和平,他们将会采取怎样的行动…… 《超時空要塞》是1982至83年间播出的一部电视动画,本片在其基础上精简改编而成。
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