上杉景虎原名北条氏秀，是北条氏康的七子，在北条与上杉家结盟之后被家人送到上杉家做上杉谦信的义子，被赐名为上杉景虎。1578年，上杉谦信病死，景虎和上杉谦信的另一个养子上杉景胜之间展开了争夺家督继承权的战争史称“御馆之乱”。景虎本来处于优势，但情况发生了大逆转，景虎被逼自杀。 本故事中的主人公仰木高耶就是上杉景虎的转世，不过他已经完全没有了前世的记忆，直到上杉家的家臣直江信纲的出现，才发现身边的一切都与平时不同了。昔日与上杉景虎志处同一时代的人慢慢地向着仰木高耶靠拢过来，离奇的事件也一再地发生。武田信玄将自己的魂魄附在了高耶的朋友成田让的身上，虽然直江想用手环将其封印却没有起作用。为了救让，高耶去了松本城，一群武田信玄手下的骸骨武士出现在高耶的周围，虽然在直江的帮助下击败了它们并成功地救出了让，但麻烦的事情才刚刚开始……
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