近未来,当年繁华喧嚣的东京因核电站事件变成了死城,被成为世界第一大幽灵城市。然而即便在这座死寂的废墟中,仍有幸村者盘桓于此。20XX年,三名没有配备任何防辐射装备的少女出现在旧东京。她们名叫成濑荆(户松遥 配音)、深作葵(花泽香菜 配音)、野村妙子(明坂聪美 配音),是通过遗传基因改造生下来天生具有抗辐射能力的少女,统称“COPPELION”。在防卫大学附属特别工科学校教官三岛鬼兵陆佐(小山力也 配音)的指挥下,少女们四处搜索幸存者,前所未有的经历以及感受接踵而至,少女们的心中作何感想……本片根据井上智德的漫画原作改编,COPPELION少女们的姓氏均来自日本著名导演,普通人姓氏则来自日本的文豪作家。
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