一种名为“卡巴内”的钢铁怪物忽然出现在人类社会之中,造成了巨大的恐慌。被它们袭击过的人类,会拥有起死回生之能力,并且亦开始袭击同类。在怪物猛烈的进攻之下,人类社会很快就四分五裂,幸存下来的人们建立起了城寨“驿”,靠着彼此互通有无苟且残存。驿与驿之间,唯一能够通行的交通工具是装甲蒸汽机车。生驹(畠中祐 配音)是一名以蒸汽锻冶为业的热血少年,他发明了武器“贯筒”,希望能够为消灭卡巴内贡献出自己的力量。一次偶然中,生驹邂逅了神秘的少女(千本木彩花 配音),与此同时,一场暴走事件的发生使得生驹所生活的显金驿彻底沦陷,卡巴内不断的产生着,生驹并没有选择逃跑,而是拿起武器,挺身而出。
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